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Journal Articles

88. Tse, P. U. (2020). Abutting Objects Warp the Three-Dimensional Curvature of Modally Completing Surfaces. Iperception. 11(2):2041669520903554. doi: 10.1177/2041669520903554.

87. Özkan M, Tse PU, Cavanagh P. (2020). Pop-out for illusory rather than veridical trajectories with double-drift stimuli. Atten Percept Psychophys. doi: 10.3758/s13414-020-02035-w.

86. Hui, J., Wang, Y., Zhang, P., Tse, P. U., Cavanagh, P. (2020). Apparent motion is computed in perceptual coordinates. i-Perception, 11(4), 1–10.

85. Cavanagh P., Tse, P. U. (2019) The vector combination underlying the double-drift illusion is based on motion in world coordinates: Evidence from smooth pursuit. Journal of Vision.

84. Liu, S., Yu Q., Tse, P. U., Cavanagh, P. (2019) Neural Correlates of the Conscious Perception of Visual Location Lie Outside Visual Cortex. Current Biology.

83. Reavis, EA, Frank, S, Tse, PU (2018) Learning efficient visual search for stimuli containing diagnostic spatial configurations and color-shape conjunctions. Attention, Perception & Psychophysics. In press.

82. Liu, S., Tse, P. U., Cavanagh, P. (2018). Meridian Interference reveals Neural Locus of Motion-induced Position Shifts. Journal of Neurophyiology.

81. Schlegel, A., Vance, B., Alxander, P., and Tse, P. U. (2018). Multivariate directed connectivity analysis (MDCA) enables decoding of directed interactions in neural and social networks.

80. Tse, P. U. (2017). Dynamic volume completion and deformation. i-Perception 8 (6), 2041669517740368.

79. Tse, P. U. (2017). Volume completion between contour fragments at discrete depths. i-Perception 8 (6),2041669517747001.

78. Sun, L. W., Hartstein, K. C., Frank, S. M., Hassan,W. and Tse, P. U. (2017). Back from the future: Volitional postdiction of perceived apparent motion direction. Vision Research. pii: S0042-6989(17)30169-4. doi: 10.1016/j.visres.2017.09.001.

77. D'Andrea-Penna, G M., Frank, S. M., Heatherton, T. F. and Tse, P. U. (2017). Distracting Tracking: Interactions between Negative Emotion and Attentional Load in Multiple Object Tracking. Emotion. Jun 1. doi: 10.1037/emo0000329.

76. Tse, P. U. (2017). Modal and amodal completion in the artwork of Coles Phillips. Perception. 46(9):1011-1013. doi: 10.1177/0301006617699666.

75. Kohler, P., Cavanagh, P. and Tse, P.U. (2017). Motion-Induced Position Shifts Activate Early Visual Cortex. Frontiers in Neuroscience. doi: 10.3389/fnins.2017.00168.

74. Frank, S.M., Greenlee, M.W. and Tse, P.U. (2017). Long time no see: enduring behavioral and neuronal changes in perceptual learning of motion trajectories three years after training. Cerebral Cortex. Feb 23:1-12. doi: 10.1093/cercor/bhx039.

73. Frank, S.M., Sun, L., Forster, L., Tse, P.U. and Greenlee, M.W. (2016). Cross-modal attention effects in vestibular cortex during attentive tracking of moving objects. J Neuroscience, pii: 2480-16.

72. Schlegel, A., Konuthula, D., Alexander, P., Blackwood, E. and Tse, P. U. (2016). Fundamentally distributed information processing integrates the motor network into the mental workspace during mental rotation. J Cogn Neuroscience, 28(8):1139-51. doi: 10.1162/jocn_a_00965.

71. Frank, S., Reavis, E., Tse, P. U. and Greenlee, M. W. (2016). Pre-training cortical thickness predicts subsequent perceptual learning rate in a visual search task. Cerebral Cortex. pii: bhu309

70. Reavis, E. A.., Frank, S., Greenlee, M. and Tse, P. U. (2016). Neural correlates of context-dependent feature-conjunction learning in visual search tasks. Human Brain Mapping, doi: 10.1002/hbm.23176.

69. Alexander, P., Schlegel, A. Sinnott-Armstrong, W., Roskies, A., Wheatley, T. and Tse, P. U. (2016). Readiness potentials driven by non-motoric processes. Consciousness & Cognition, 39:38-47. doi: 10.1016/j.concog.2015.11.011.

68. Schlegel, A., Alexander, P. and Tse, P. U. (2016). Information processing in the mental workspace is fundamentally distributed. Journal of Cognitive Neuroscience. J Cogn Neurosci, 28(2):295-307. doi: 10.1162/jocn_a_00899.

67. McCarthy, J. D., Kohler, P. J., Tse, P. U. and Caplovitz, G. P. (2015). Extrastriate visual areas integrate form features over space and time to construct representations of stationary and rigidly rotating objects. Journal of Cognitive Neuroscience. July 30, 1-17. PMID 26226075.

66. Kohler, P. J., Cavanagh, P. and Tse, P. U. (2015). Motion-induced position shifts are influenced by global motion, but dominated by component motion. Vision Research, 110(Pt A): 93-9. doi: 10.1016/j.visres.2015.03.003.

65. Mastropasqua T., Tse, P. U., and Turatto, M. (2015). Learning of monocular information facilitates breakthrough to awareness during interocular suppression. Atten Percept Psychophys. 77(3):790-803. doi: 10.3758/s13414-015-0839-z.

64. Reavis, E.A., Frank, S.M. and Tse, P.U. (2015). Caudate nucleus reactivity predicts perceptual learning rate for visual feature conjunctions. Neuroimage; 110C:171-181. doi: 10.1016/j.neuroimage.2015.01.051.

63. Schlegel, A., Alexander, P., Sinnott-Armstrong, W., Roskies, A., Tse, P.U., Wheatley, T. (2015). Hypnotizing Libet: Readiness potentials with non-conscious volition. Consciousness and Cognition;33C:196-203. doi: 10.1016/j.concog.2015.01.002.

62. Schlegel, AS, Alexander, P, Fogelson, SV, Li, X, Lu, Z, Kohler, PJ, Riley, E, Tse, P.U, and Meng, M. (2015). The artist emerges: Visual art learning alters neural structure and function. Neuroimage, 105:440-51. doi: 10.1016/j.neuroimage.2014.11.014.

61. Fogelson SV, Kohler PJ, Miller KJ, Granger R, Tse PU. (2014). Unconscious neural processing differs with method used to render stimuli invisible. Front Psychol. 5:601. doi: 10.3389/fpsyg.2014.00601

60. Kohler PJ, Caplovitz GP, Tse PU. (2014). The global slowdown effect: why does perceptual grouping reduce perceived speed? Atten Percept Psychophys. 76(3):780-92. doi: 10.3758/s13414-013-0607-x.

59. Frank, S., Reavis, E. A., Tse, P. U. and Greenlee, M. (2014). Neural mechanisms of feature conjunction learning: enduring changes in occipital cortex after a week of training. Human Brain Mapping. doi: 10.1002/hbm.22245

58. Schlegel, A., Kohler, P., Fogelson, S., Alexander, S. and Tse, P. U. (2013). Network structure and dynamics of the mental workspace. Proceedings of the National Academy of Sciences, 110(40):16277-82. doi: 10.1073/pnas.1311149110.

57. Reavis, E. A., Kohler, P. J., Caplovitz, G. P., Wheatley, T. P. and Tse, P. U. (2013). Effects of attention on visual experience during monocular rivalry. Vision Research. doi:pii: S0042-6989(13)00057-6. 10.1016/j.visres.2013.03.002.

56. Schlegel, A., Alexander, P., Sinnott-Armstrong, W., Roskies, A., Tse, P. U., and, Wheatley, T. (2013). Barking up the wrong free: readiness potentials reflect processes independent of conscious will. Experimental Brain Research. 229(3):329-35. doi: 10.1007/s00221-013-3479-3.

55. Kohler PJ, Fogelson SV, Reavis EA, Meng M, Guntupalli JS, Hanke M, Halchenko YO, Connolly AC, Haxby JV, Tse PU. (2013). Pattern classification precedes region-average hemodynamic response in early visual cortex. Neuroimage. 78:249-60. doi: 10.1016/j.neuroimage.2013.04.019.

54. Kosovicheva, A. A., Maus, G. W., Anstis, S., Cavanagh, P., Tse, P. U., Whitney, D. (2012). The motion-induced shift in the perceived location of a grating also shifts its aftereffect. Journal of Vision, 12(8) doi 10.1167/12.8.7.

53. Schlegel AA, Rudelson JJ, Tse P. U. (2012). White matter structure changes as adults learn a second language. Journal of Cognitive Neuroscience, 24(8):1664-70.

52. Porter, K. B., Caplovitz, G. P., Kohler, P. J., Ackerman, C. M. and Tse, P. U. (2011). Rotational and translational motion interact independently with form. Vision Res., 51(23-24):2478-87.

51. Tse, P. U., Whitney, D., Anstis, S. and Cavanagh, P. (2011). Voluntary attention modulates motion-induced mislocalization. J. of Vision, 11(3):12

50. Caplovitz, G. P. and Tse, P. U. (2010). Extrastriate cortical activity reflects segmentation of motion into independent sources. Neuropsychologia. 48(9):2699-708.

49. Kohler, P. K., Caplovitz, G. P., Hsieh, P.-J., Sun, J., and Tse, P. U. (2010). Motion fading is driven by perceived, not actual angular velocity. Vision Research, 50(11):1086-94.

48. Hsieh, P.-J. and Tse, P. U. (2010). BOLD signal in both ipsilateral and contralateral retinotopic cortex modulates with perceptual fading. PLOS One, 5(3):e9638.

47. Hsieh, P.-J. and Tse, P. U. (2010). 'Brain-reading' of perceived colors reveals a feature mixing mechanism underlying perceptual filling-in in cortical area V1. Human Brain Mapping. Epub Jan. 19, 2010.

46. Tse, P. U., Baumgartner, F. J., and Greenlee, M. W. (2010). Event-related functional MRI of cortical activity evoked by microsaccades, small visually-guided saccades, and eyeblinks in human visual cortex. Neuroimage, 49, 805-816.

45. Hsieh, P.-J. and Tse, P. U. (2009). Microsaccade rate varies with subjective visibility during motion-induced blindness. PLOS One. 4, 4, e5163.

44. Hsieh, P.-J. and Tse, P. U. (2009). Motion fading and the motion after-effect share a common process of neural adaptation. Attention, Perception & Psychophysics. 71(4),724-733.

43. Tse, P. U., Caplovitz, G. P., and Hsieh, P.-J. (2009). Corrigendum to ‘‘Microsaccade directions do not predict directionality of illusory brightness changes of overlapping transparent surfaces” [Vision Research 46 (2006) 3823–3830]. Vision Research 49, 790, e1-e7.

42. Kohler, P. K., Caplovitz, G. P., and and Tse, P. U. (2009). The whole moves less than the spin of its parts. Attention, Perception & Psychophysics. 71(4), 675-679.

41. Hsieh, P.-J. and Tse, P. U. (2009). Feature mixing rather than feature replacement during perceptual filling-in. Vision Research. 49(4):439-50.

40. Caplovitz, G. P., Paymer, N., and Tse, P. U. (2008). The Drifting Edge Illusion: A stationary edge abutting an oriented drifting grating appears to move because of the 'other aperture problem.' Vision Research, 48(22):2403-14.

39. Chiao, J. Y., Adams, R. B., Tse, P. U., Lowenthal, W. T., Richeson, J. A. and Ambady, N. (2008). Knowing who's boss: fMRI and ERP investigations of social dominance perception. Group Processes and Intergroup Relations, special issue on Social Neuroscience, April, 2008, 11(2) 201-214.

38. Caplovitz, G. P., Barroso, D. J., Hsieh, P.-J. and Tse, P. U. (2007). fMRI reveals that neuronal feedback to ventral retinotopic cortex underlies perceptual grouping by temporal synchrony. Human Brain Mapping, 29(6):651-61

37. Tse, P. U. and Hsieh, P.-J. (2007). Component and intrinsic motion integrate in ‘dancing bar’ illusion. Biological Cybernetics, 96(1):1-8.

36. Caplovitz, G. P. and Tse, P. U. (2007). Rotating Dotted Ellipses: Motion perception driven by grouped figural rather than local dot motion signals. Vision Research, 47(15):1979-91.

35. Troncoso, X. G., Tse, P. U., Macknik, S. L., Caplovitz,, G. P., Hsieh, P.-J. Schlegel, A. A., Martinez-Conde, S. (2007). BOLD activation varies parametrically with corner angle in all human retinotopic areas. Perception, 36(6):808-20.

34. Hsieh, P.-J. and Tse, P. U. (2007). Grouping inhibits motion fading by giving rise to virtual trackable features. Journal of Experimental Psychology: Human Perception and Performance, 33(1):57-63.

33. Caplovitz, G. P. and Tse, P. U. (2007). V3A processes contour curvature as a trackable feature for the perception of rotational motion. Cerebral Cortex. 17,1179-1189.

32. Tse, P. U. and Hsieh, P.-J. (2006). The infinite regress illusion reveals faulty integration of local and global motion signals. Vision Research, 46(22):3881-5.
View Demo Here

31. Caplovitz, G. P. and Tse, P. U. (2006). The bar-cross-ellipse illusion: alternating percepts of rigid and nonrigid motion based on contour ownership and trackable feature assignment. Perception, 35(7):993-7.
View Demo Here

30. Tse, P. U., Caplovitz, G. P., and Hsieh, P.-J. (2006). Microsaccade directions do not predict directionality of illusory brightness changes of overlapping transparent surfaces. Vision Research, 46(22):3823-30. Epub 2006 Aug 24.

29. Hsieh, P.-J., Caplovitz, G. P., and Tse, P. U. (2006). Bistable illusory rebound motion: Event-related functional magnetic resonance imaging of perceptual states and switches. Neuroimage, 32(2):728-39.

28. Caplovitz, G. P. and Tse, P. U. (2006). Mechanisms underlying the perceived angular velocity of a rigidly rotating object. Vision Research, 46(18):2877-93.

27. Tse, P. U. (2006). Neural correlates of transformational apparent motion. Neuroimage, 31(2):766-73.

26. Hsieh, P.-J. and Tse, P. U. (2006). Illusory color mixing upon perceptual fading and filling-in does not result in ‘forbidden colors.’ Vision Research, 46(14):2251-8.

25. Hsieh, P.-J. and Tse, P. U. (2006). Stimulus factors affecting Illusory Rebound Motion. Vision Research, 46(12):1924-33.

24. Hsieh, P.-J., Caplovitz, G. P., and Tse, P. U. (2006). Illusory motion induced by the offset of stationary luminance-defined gradients. Vision Research, 46(6-7):970-8.

23. Tse, P. U., Martinez-Conde, S., Schlegel, A., and Macknik, S. (2005). Visibility and visual masking of simple targets are confined to areas in the occipital cortex beyond human V1/V2. Proceedings of the National Academy of Sciences, 102, 47, 17178-17183.

22. Eagleman, D. M., Tse, P. U., Janssen, P., Nobre, A. C., Buonomano, D., and Holcombe, A. O. (2005). Time and the brain: how subjective time relates to neural time. Journal of Neuroscience, 25(45):10369-71.

21. Hsieh, P.-J., Caplovitz, G. P., and Tse, P. U. (2005). Illusory Rebound Motion and the motion continuity heuristic. Vision Research, 45(23):2972-85.

20. Tse, P. U. (2005). Voluntary attention modulates the brightness of overlapping transparent surfaces. Vision Research, 45(9):1095-8.

19. Tse, P. U. (2004). Unser Ziel muss eine Gestalt-Neurowissenschaft sein. Gestalt Theory, 287-292.

18. Tse, P. U., Rivest, J., Intriligator, J. and Cavanagh, P. (2004). Attention and the subjective expansion of time. Perception & Psychophysics, 66(7), 1171-1189.

17. Tse, P. U., Sheinberg, D. L., and Logothetis, N. K. (2004). The distribution of microsaccade directions need not reveal the location of attention. Psychological Science, 15(10):708-10.

16. Tse, P. U. (2004). Mapping visual attention with change blindness: new directions for a new method. Cognitive Science, 28, 2, 241-258.

15. Tse, P. U. (2003). If vision is 'veridical hallucination', what keeps it veridical? Commentary on “Gestalt isomorphism and the primacy of subjective conscious experience: a Gestalt Bubble model” by Steven Lehar. Behavioral and Brain Sciences, 26(4):375-408.

14. Tse, P. U., Sheinberg, D. L., and Logothetis, N. K. (2003). Attentional enhancement opposite a peripheral flash revealed by change blindness. Psychological Science, 14, 2, 1-8.

13. Tse, P. U., Sheinberg, D. L., and Logothetis, N. K. (2002). Fixational eye movements are not affected by abrupt onsets that capture attention. Vision Research, 42, 1663-1669.

12. Tse, P. U. and Logothetis, N. K. (2002). The duration of 3-D form analysis in transformational apparent motion. Perception & Psychophysics, 64(2), 244-265.

11. Tse, P. U. (2002). A contour propagation account of surface filling-in and volume formation. Psychological Review, 109, 1, 91-115.

10. Kristjansson, A. and Tse, P. U. (2001). Curvature discontinuities are cues for rapid shape analysis. Perception & Psychophysics, 63, 390-403.

9. Tse, P. U. and Cavanagh, P. (2000). Chinese and Americans see opposite apparent motions in a Chinese character. Cognition, 74, B27-B32.

8. Tse, P. U. (2000). The sawtooth illusion. Perception. 29(7), 874-876.

7. Holcombe, A. O., Intriligator, J., & Tse, P. U. (2000). The spokes illusion originates at an early motion processing stage. Perception & Psychophysics, 62(8), 1619-1624.

6. Albert, M. and Tse, P. U. (2000). The role of surface attachment in perceived volumetric shape. Perception, 29, 409-420.

5. Tse, P. U. (1999). Volume completion. Cognitive Psychology, 39, 37-68.

4. Tse, P. U. (1999). Complete mergeability and amodal completion. Acta Psychologica, 102, 165-201.

3. Holcombe, A. O., Macknik, S. L., Intriligator, J., Seiffert,A. E., and Tse, P. U. (1999). Wakes and spokes: New motion-induced brightness illusions. Perception, 28, 10, 1231-1242.

2. Tse, P. and Albert, M. (1998). Amodal Completion in the absence of image tangent discontinuities. Perception, 27, 455-464.

1. Tse, P. (1998). Illusory volumes from conformation. Perception, 27, 8, 977-994.

Popular Articles

Tse, P. U. (2013). Free Will Unleashed in New Scientist, 8 June 2013, pp 28-29


Tse, P. U. (2017). Two types of libertarian free will are realized in the human brain. In Gregg Caruso “Neuroexistentialism”. Oxford University Press.

Alexander, P., Schlegel, A., Sinnott-Armstrong, W., Roskies, A., Tse, P. U., & Wheatley, T. (2014). Dissecting the Readiness Potential: An Investigation of the Relationship between Readiness Potentials, Conscious Willing, and Action. In Mele (Ed.). Surrounding Free Will: Philosophy, Psychology, Neuroscience; Oxford University Press

Blair, C. D., Tse, P. U. and Caplovitz, G. P. (2013). Interactions of form and motion in the perception of moving objects. Oxford Handbook of Perceptual Organization. ed. Johan Wagemans. Oxford Press. U.K.

Tse, P. U. and Palmer, S. E (2013). Visual perceptual organization. Ed. By K. Ochsner and S. Kosslyn. Oxford Handbook. Submitted. Oxford U. Press.

Tse, P. U., Reavis, E. A., Kohler, P. J., Caplovitz, G. P. and Wheatley, T. (2013). How attention can alter appearances. Wiley Handbook on Perception of Appearances. ed. by Liliana Albertazzi.

Tse, P. U. and Palmer, S. E (2012). Visual object processing. Wiley Handbook on Perception. Ed. By Healy, Wiener and Proctor. Oxford U. Press.

Attention causes the subjective expansion of time. Kia Nobre ed., Oxford University Press.

Greenlee, M. W. and Tse, P. U. (2008). Functional neuroanatomy of the human visual system: a review of functional MRI studies. In: ‘Essentials in Ophthalmology,‘ Springer Verlag, Germany.

Tse, P. U. and Caplovitz, G. P. (2006). Contour discontinuities subserve two types of form analysis that underlie motion processing. In: ‘Progress in Brain Research: Visual Perception Part I, Fundamentals of Vison: Low and Mid-Level processes in perception.’ Elsevier. 154:271-92.

Tse, P. (2006). How the evolution of symbolic cognition transformed human morality. In: ‘Moral Psychology, Volume 1: The Evolution of Morality.’ Oxford University Press. Edited by Sinnott-Armstrong.

Tse, P. U. and Hughes, H. C. (2004). Visual Pattern and Form Perception. In: ‘The Encyclopedia of Neuroscience.’ Adelman, G. and Smith, B. (Eds.). Elsevier.

Tse, P., Cavanagh, P., and Nakayama, K. (1998). The role of parsing in high-level motion processing. In: ‘High-level motion processing - Computational, neurobiological and psychophysical perspectives.’ Watanabe, T. (Ed.). MIT Press, pp. 249-266.

Op Eds

Tse: Liberating Minds to the Core

Psychology and Neuroscience Books

The neural basis of free will: criterial causation.

Progress in Brain Research: Visual Perception Part 1: Fundamentals of Vision, Low and Mid-level processes in Perception’ Edited by S. Martinez-Conde, S.L. Macknik, L.M. Martinez, J.-M. Alonso and P.U. Tse. This is vol. 154 of the book series.

‘Progress in Brain Research: Visual Perception Part 2: Fundamentals of Aareness, Multi-Sensory Integration and High-Order Perception’ Edited by S. Martinez-Conde, S.L. Macknik, L.M. Martinez, J.-M. Alonso and P.U. Tse. This is vol. 155 of the book series.